Natural selection is the process by which organisms adapt to their environment over generations. This fundamental concept in biology explains how populations evolve, with traits that enhance survival and reproduction becoming more common over time.
Darwin's observations during his voyage on the HMS Beagle led to the development of this theory. Evidence from fossils, comparative anatomy, and molecular biology supports natural selection as the driving force behind the diversity of life on Earth.
Natural selection process by which populations become adapted to their environment over the course of generations
Adaptation a trait that increases an organism's fitness and arises through natural selection
Can be morphological (physical), physiological, or behavioral
Fitness the ability to survive and reproduce in a specific environment
Variation the presence of differences among individuals in a population
Variation is necessary for natural selection to occur
Inheritance the passing of traits from parents to offspring
Differential reproduction occurs when individuals with certain traits leave more offspring than individuals without those traits
Speciation the formation of new species as a result of evolution by natural selection
Historical Context and Darwin's Contributions
Charles Darwin developed the theory of evolution by natural selection in the mid-19th century
Darwin's voyage on the HMS Beagle exposed him to diverse flora and fauna and geological evidence that influenced his thinking
Darwin observed that organisms seemed well-suited to their environments and that similar species often differed across geographic regions
Alfred Russel Wallace independently developed a theory of evolution by natural selection and corresponded with Darwin
Darwin's book "On the Origin of Species" (1859) outlined his evidence for evolution and proposed natural selection as the mechanism
The book was controversial at the time but eventually gained widespread acceptance in the scientific community
Darwin's theory built upon earlier ideas, such as Lamarck's concept of the inheritance of acquired characteristics
Modern evolutionary theory synthesizes Darwin's ideas with Mendelian genetics and other scientific discoveries
Mechanisms of Natural Selection
Overproduction more offspring are produced than can survive due to limited resources
Genetic variation exists within populations for traits that affect survival and reproduction
Variation arises through mutation, recombination during sexual reproduction, and gene flow between populations
Differential survival and reproduction individuals with traits that are advantageous in a given environment tend to survive and reproduce more than those without those traits
Over time, this leads to a change in the frequency of alleles in the population
Heredity traits that confer an advantage are passed down to offspring
Accumulation of changes over many generations, the advantageous traits become more common in the population
Adaptation to the environment results from the gradual accumulation of beneficial traits
Speciation can occur when populations become reproductively isolated and diverge due to different selective pressures
Evidence for Natural Selection
Fossil record shows gradual changes in organisms over time and the existence of extinct species
Transitional fossils demonstrate the evolution of one species into another
Comparative anatomy reveals structural similarities among related species, suggesting common ancestry
Homologous structures (e.g., human arm, bat wing, whale flipper) have similar underlying anatomy due to shared ancestry
Analogous structures (e.g., bird wing, insect wing) have similar functions but different underlying anatomy due to convergent evolution
Embryology early developmental stages of different species show similarities, supporting common ancestry
Molecular biology DNA and protein sequences show similarities among related species
The degree of similarity reflects the relatedness of species
Biogeography the distribution of species across geographic regions reflects their evolutionary history
Island species often resemble nearby mainland species but with unique adaptations
Observed instances of natural selection (e.g., antibiotic resistance in bacteria, beak shape changes in Galápagos finches)
Types of Selection
Directional selection favors individuals at one extreme of a trait distribution
Leads to a shift in the population's mean trait value over time
Stabilizing selection favors individuals with intermediate trait values
Reduces variation in the population and maintains the mean trait value
Disruptive selection favors individuals at both extremes of a trait distribution
Can lead to the formation of distinct subpopulations or even new species
Sexual selection occurs when traits that enhance mating success are favored
Can result in the evolution of exaggerated traits (e.g., peacock tail)
Artificial selection human-driven selection for desired traits in domesticated plants and animals
Demonstrates the power of selection to alter populations
Adaptation and Fitness
Adaptation an inherited trait that enhances an organism's fitness in a specific environment
Fitness the ability to survive and reproduce in a given environment
Measured by an individual's relative contribution to the next generation
Adaptations can take many forms, such as morphological, physiological, or behavioral traits